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Trilobites A to Z
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22 September 2026

Trilobites are among the most remarkable lifeforms ever to exist. First emerging 521 million years ago, these organisms—animals with highly developed eyes, sturdy walking legs, and protective outer shells—evolved into more than 25,000 scientifically recognized species. During their 270 million years of inhabiting the ancient seas, trilobites proliferated into an ever-more-fanciful cast of characters, from graceful Cambrian olenellids to robust Ordovician asaphids to spine-covered Devonian phacopids. Their fossil remains, which today can be found on every continent, testify to the astonishing breadth and scope of trilobite heterogeneity.
Trilobites A to Z is the ultimate reference guide to these incredible invertebrates. It features reader-friendly, concise yet information-packed introductions to nearly 1,500 trilobite species, each accompanied by a full-color photograph. Grouped according to periods within the Paleozoic era, the book illuminates the similarities and differences among trilobite specimens from around the globe, bringing to life the complexity and beauty of these long-extinct creatures in all their variety. Andy Secher—one of the world’s leading trilobite collectors—also offers practical guidance for readers seeking to identify specimens of their own. Comprehensive and entertainingly written, this essential encyclopedia will inform and delight trilobite enthusiasts and collectors as well as students and scholars of invertebrate paleontology.
— Martin Shugar, paleontology field associate, American Museum of Natural History
Andy Secher's alphabetical romp through the Paleozoic world of trilobites is a stunning visual testament to their enduring appeal—a fossil feast for the eyes!
— David Rudkin, former assistant curator of invertebrate paleontology, Royal Ontario Museum
Trilobites A to Z is a comprehensive reference for the incredible diversity of trilobites in a format useful to collectors and scientists. Secher’s skill in describing complex scientific principles like taxonomy, evolution, and geology in an accessible style offers a clever exploration of these captivating animals for anyone who appreciates their beauty and utility.
— Susan Butts, Yale Peabody Museum
Andy Secher has been a field associate in paleontology at the American Museum of Natural History for more than two decades. He curates one of the world’s largest private trilobite collections. Secher is the author of Travels with Trilobites: Adventures in the Paleozoic (Columbia, 2022) and The Trilobite Collector’s Guide (Columbia, 2024), and he was the longtime editor of the rock music magazine Hit Parader.
David Rudkin is the former assistant curator of invertebrate paleontology at the Royal Ontario Museum.
Greg Edgecombe is a merit researcher in the Department of Earth Sciences at the Natural History Museum, London.
Foreword: The Timeless Trilobite, by David Rudkin
Foreword: Some Reasons to Ponder Peachella, by Greg Edgecombe
The Story Behind the Photographs
Introduction: Trilobites A to Z
Why Metric Measurements?
The Paleozoic Era (541–252 Million Years Ago)
The Cambrian Period (541–485 Million Years Ago)
The Ordovician Period (485–444 Million Years Ago)
The Silurian Period (444–419 Million Years Ago)
The Devonian Period (419–359 Million Years Ago)
The Mississippian Period (359–323 Million Years Ago)
The Pennsylvanian Period (323–299 Million Years Ago)
The Permian Period (299–252 Million Years Ago)
Final Thought
Special Thanks
Glossary: Trilobite Terminology
Index